Once again, the Imperial Fighting Troupe has been called out to do battle with vicious demons that are attacking the new capital city, Tokyo. Unfortunately, something stinks in suburbia, and an insidious American company called Douglas-Stewart has entered the fray, fighting back the demons with mechanized robots more powerful than the Troupe's! Douglas-Stewart's champion robots make life miserable for Sakura and her plucky gang of heroines, who are on indefinite stand-by status. When Maria investigates Douglas-Stewart, all hell breaks loose, and it's up to the Imperial Fighting Troupe to crack the mystery and regain their glory!
《樱花大战活动写真》的故事发生在大神前往巴黎后的一段时间,而这也正是大家非常想了解的一段剧情“大神在巴黎时,樱在干什么?”。剧情很好的衔接与弥补了2代、3代和4代之间的空白。
与以往游戏和动画不同,观众们将首次看到没有大神登场的《樱花大战》故事。剧场动画的时间恰好设定在大神一郎赴法国修行的时候,而且是在日本的花组队员前往法国“探亲”之前。当时远在巴黎的大神正在欧洲着手组建巴黎华击团,留在帝都的花组队员们也迎来了原星组的队长拉切特.阿尔泰尔。正当信奉合理主义和个人主义的拉切特和有些大而化之的花组队员们较劲时,神秘的降魔军团再次威胁帝都。针对花组连续多次在战斗中失态的状况,竟然出现了解散花组的舆论呼声,队长不在、内外交困的花组遭遇了前所未有的危机......