Ryotaro Nogami is a young man with a lot of bad luck. One day, he finds a strange pass and things got stranger from a mysterious girl and a large time-traveling train to being possessed by an entity called an Imagin, beings from an alternate future whose kind are attempting to change the past. Though slightly confused about the nature of the crisis, Ryotaro, along with the aid of the hot-headed, violent Imagin, dubbed Momotaros, becomes Kamen Rider Den-O, traveling to different times on the Den-Liner to battle the evil Imagin to prevent them from altering the past to affect the present and future. During his adventure, Ryotaro is joined by other Imagin who aid him as well; the lying, manipulating and womanizing Urataros, the herculean (and narcoleptic) Kintaros, and the childish yet powerful Ryutaros. He later meets the mysterious Yuto Sakurai and his bumbling Imagin partner Deneb. Yuto is not only Kamen Rider Zeronos but is the younger incarnation of Ryotaro's older sister Airi's fiance, Sakurai, who mysteriously disappeared and is tied to the mysteries revolving the Imagin and a person known as the Junction Point.
18岁的少年，主角野上良太郎虽拥有普通平凡的外表，但却是史上最多恶运缠身的男子！某天。良太郎进入了不可思议的路径，并遇上欲凭衣到他身上的可怕魔神 Momotarosu，幸得神秘的女子Haha及时阻止，而且良太郎更从她手上得到Rider Pass，获得变身成为电王的资格！良太郎之后更将魔神Momotarosu的邪恶之心封印，保留其英雄格~化作电王的正义力量！新的历史即将开始，2007年，出现了依凭实现人类的欲望，代价是该人类的过去，来自未来的侵略者-异魔神。其目的是想夺取对方的回忆回到过去。改变历史从而改变未来。
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